Deletion of the gene encoding the reductase component of 3-ketosteroid 9α-hydroxylase inRhodococcus equiUSA-18 disrupts sterol catabolism, leading to the accumulation of 3-oxo-23,24-bisnorchola-1,4-dien-22-oic acid and 1,4-androstadiene-3,17-dione

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